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Content text 6. READING: TULIPA (TULIPS) - Anna Mai.pdf

TULIPA (TULIPS) Tulipa (tulips) is a genus of spring-blooming perennial herbaceous bulbiferous geophytes, dying back after flowering to an underground storage bulb. Depending on the species, tulip plants can be between 10 and 70 cm (4 and 28 inches) high. Flowers: The tulip's flowers are usually large and are actinomorphic (radially symmetric) and hermaphrodite (contain both male (androecium) and female (gynoecium) characteristics), generally erect, or more rarely pendulous, and are arranged more usually as a single terminal flower, or when pluriflor as two to three (e.g. Tulipa turkestanica), but up to four, flowers on the end of a floriferous stem (scape), which is single arising from amongst the basal leaf rosette. In structure, the flower is generally cup or star shaped. As with other members of Liliaceae the perianth is undifferentiated (perigonium) and biseriate (two whorled), formed from six free (i.e. apotepalous) caducous tepals arranged into two separate whorls of three parts (trimerous) each. The two whorls represent three petals and three sepals, but are termed tepals because they are nearly identical. The tepals are usually petaloid (petal like), being brightly coloured, but each whorl may be different, or have different coloured blotches at their bases, forming darker colouration on the interior surface. The inner petals have a small, delicate cleft at the top, while the sturdier outer ones form uninterrupted ovals. Androecium: The flowers have six distinct, basifixed introrse stamens arranged in two whorls of three, which vary in length and may be glabrous or hairy. The filaments are shorter than the tepals and dilated towards their base. Gynoecium: The style is short or absent and each stigma has three distinct lobes, and the ovaries are superior, with three chambers. Fruit: The tulip's fruit is a globose or ellipsoid capsule with a leathery covering and an ellipsoid to globe shape. Each capsule contains numerous flat, disc-shaped seeds in two rows per chamber. These light to dark brown seeds have very thin seed coats and endosperm that does not normally fill the entire seed. Anna Mai https://www.facebook.com/AnnaMai.AptisSV/ 01
TULIPA (TULIPS) Leaves: Tulip stems have few leaves. Larger species tend to have multiple leaves. Plants typically have two to six leaves, some species up to 12. The tulip's leaf is cauline (born on a stem), strap-shaped, with a waxy coating, and the leaves are alternate (alternately arranged on the stem), diminishing in size the further up the stem. These fleshy blades are often bluish-green in colour. The bulbs are truncated basally and elongated towards the apex. They are covered by a protective tunic (tunicate) which can be glabrous or hairy inside. Anna Mai https://www.facebook.com/AnnaMai.AptisSV/ The "Semper Augustus" was the most expensive tulip during the 17th-century tulip mania. “The colour is white, with Carmine on a blue base, and with an unbroken flame right to the top” – wrote Nicolas van Wassenaer in 1624 after seeing the tulip in the garden of one Dr Adriaen Pauw, a director of the new East India Company. With limited specimens in existence at the time and most owned by Pauw, his refusal to sell any flowers, despite wildly escalating offers, is believed by some to have sparked the mania. Colours Tulip flowers come in a wide variety of colours, except pure blue (several tulips with "blue" in the name have a faint violet hue), and have absent nectaries. Tulip flowers are generally bereft of scent and are the coolest of floral characters. The Dutch regarded this lack of scent as a virtue, as it demonstrates the flower's chasteness. While tulips can be bred to display a wide variety of colours, black tulips have historically been difficult to achieve. The Queen of the Night tulip is as close to black as a flower gets, though it is, in fact, a dark and glossy maroonish purple - nonetheless, an effect prized by the Dutch. The first truly black tulip was bred in 1986 by a Dutch flower grower in Bovenkarspel, Netherlands. The specimen was created by cross-breeding two deep purple tulips, the Queen of the Night and Wienerwald tulips. 02
TULIPA (TULIPS) Anna Mai https://www.facebook.com/AnnaMai.AptisSV/ Tulipanin is an anthocyanin found in tulips. It is the 3-rutinoside of delphinidin. The chemical compounds named tuliposides and tulipalins can also be found in tulips and are responsible for allergies. Tulipalin A, or α-methylene-γ-butyrolactone, is a common allergen, generated by hydrolysis of the glucoside tuliposide A. It induces a dermatitis that is mostly occupational and affects tulip bulb sorters and florists who cut the stems and leaves. Tulipanin A and B are toxic to horses, cats and dogs. The colour of a tulip is formed from two pigments working in concert; a base colour that is always yellow or white, and a second laid-on anthocyanin colour. The mix of these two hues determines the visible unitary colour. The breaking of flowers occurs when a virus suppresses anthocyanin and the base colour is exposed as a streak. Phytochemistry The great majority of tulips, both species and cultivars, have no discernable scent, but a few of both are scented to a degree, and Anna Pavord describes T. Hungarica as "strongly scented" , and among cultivars, some such as "Monte Carlo" and "Brown Sugar" are "scented" , and "Creme Upstar" "fragrant". Fragrance Tulipa is a genus of the lily family, Liliaceae, once one of the largest families of monocots, but which molecular phylogenetics has reduced to a monophyletic grouping with only 15 genera. Within Liliaceae, Tulipa is placed within Lilioideae, one of three subfamilies, with two tribes. Tribe Lilieae includes seven other genera in addition to Tulipa. Taxonomy of Tulipa Clusianae (4 species) Orithyia (4 species) Tulipa (52 species) Eriostemones (16 species) Subdivision The genus, which includes about 75 species, is divided into four subgenera. 03
TULIPA (TULIPS) Anna Mai https://www.facebook.com/AnnaMai.AptisSV/ Eastern end of the tulip range from Turkmenistan on the eastern shore of the Caspian Sea to the Pamir-Alai and Tien-Shan mountains. Distribution and habitat Tulips are mainly distributed along a band corresponding to latitude 40° north, from southeast of Europe (Greece, Albania, North Macedonia, Kosovo, Southern Serbia, Bulgaria, most part of Romania, Ukraine, Russia) and Turkey in the west, through the Levant (Syria, Israel, Palestinian Territories, Lebanon and Jordan) and the Sinai Peninsula. From there it extends eastwards through Jerevan, (Armenia) and Baku (Azerbaijan) and on the eastern shore of the Caspian Sea through Turkmenistan, Bukhara, Samarkand and Tashkent (Uzbekistan), to the eastern end of the range in the Pamir-Alai and Tien-Shan mountains in Central Asia, which form the centre of diversity. Further to the east, Tulipa is found in the western Himalayas, southern Siberia, Inner Mongolia, and as far as the northwest of China. While authorities have stated that no tulips west of the Balkans are native, subsequent identification of Tulipa sylvestris subsp. australis as a native of the Iberian peninsula and adjacent North Africa shows that this may be a simplification. In addition to these regions in the west tulips have been identified in Greece, Cyprus and the Balkans. In the south, Iran marks its furthest extent, while the northern limit is Ukraine.[21] Although tulips are also throughout most of the Mediterranean and Europe, these regions do not form part of the natural distribution. Tulips were brought to Europe by travellers and merchants from Anatolia and Central Asia for cultivation, from where they escaped and naturalised (see map). For instance, less than half of those species found in Turkey are actually native. These have been referred to as neo-tulipae. Tulips are indigenous to mountainous areas with temperate climates, where they are a common element of steppe and winter-rain Mediterranean vegetation. They thrive in climates with long, cool springs and dry summers. Tulips are most commonly found in meadows, steppes and chaparral, but also introduced in fields, orchards, roadsides and abandoned gardens. 04

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